About the Authors

Christopher Wostenberg

Affiliation Department of Chemistry and Center for RNA Molecular Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Jeffrey W. Lary

Affiliation National Analytical Ultracentrifugation Facility, University of Connecticut, Storrs, Connecticut, United States of America

Debashish Sahu

Affiliation Department of Chemistry and Center for RNA Molecular Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Roderico Acevedo

Affiliation Department of Chemistry and Center for RNA Molecular Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Kaycee A. Quarles

Affiliation Department of Chemistry and Center for RNA Molecular Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

James L. Cole

Affiliations National Analytical Ultracentrifugation Facility, University of Connecticut, Storrs, Connecticut, United States of America, Department of Molecular and Cell Biology and Department of Chemistry, University of Connecticut, Storrs, Connecticut, United States of America

Scott A. Showalter

sas76@psu.edu

Affiliation Department of Chemistry and Center for RNA Molecular Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CW DS JLC SAS. Performed the experiments: CW JWL DS RA KAQ JLC. Analyzed the data: CW JWL JLC DS RA KAQ SAS. Contributed reagents/materials/analysis tools: CW JWL JLC DS RA KAQ SAS. Wrote the paper: CW JLC SAS.