About the Authors

Karen Staines

Affiliation Avian Viral Diseases Programme, The Pirbright Institute, Compton Laboratory, Newbury, Berkshire, United Kingdom

John R. Young

Affiliation Avian Viral Diseases Programme, The Pirbright Institute, Compton Laboratory, Newbury, Berkshire, United Kingdom

Colin Butter

colin.butter@avianimmunology.org

Affiliation Avian Viral Diseases Programme, The Pirbright Institute, Compton Laboratory, Newbury, Berkshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KS JRY CB. Performed the experiments: KS CB. Analyzed the data: KS JRY CB. Contributed reagents/materials/analysis tools: JRY. Wrote the paper: KS JRY CB.