About the Authors

Richard C. Krolewski

Contributed equally to this work with: Richard C. Krolewski, Adam Packard

Affiliations Department of Anatomy and Cellular Biology, Tufts University School of Medicine, Boston, Massachusetts, United States of America, Program in Cellular, Molecular, and Developmental Biology, Sackler School of Graduate Biomedical Sciences, Tufts University School of Medicine, Boston, Massachusetts, United States of America

Adam Packard

Contributed equally to this work with: Richard C. Krolewski, Adam Packard

Affiliations Department of Anatomy and Cellular Biology, Tufts University School of Medicine, Boston, Massachusetts, United States of America, Program in Cellular, Molecular, and Developmental Biology, Sackler School of Graduate Biomedical Sciences, Tufts University School of Medicine, Boston, Massachusetts, United States of America

Woochan Jang

Affiliation Department of Anatomy and Cellular Biology, Tufts University School of Medicine, Boston, Massachusetts, United States of America

Hendrik Wildner

Affiliation Institute of Pharmacology and Toxicology, University of Zürich, Zürich, Switzerland

James E. Schwob

jim.schwob@tufts.edu

Affiliation Department of Anatomy and Cellular Biology, Tufts University School of Medicine, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RCK AP WJ JES. Performed the experiments: RCK AP WJ. Analyzed the data: RCK AP WJ. Contributed reagents/materials/analysis tools: HW. Wrote the paper: RCK AP WJ JES.