About the Authors

Sayuri Yamazaki

Contributed equally to this work with: Sayuri Yamazaki, Akira Maruyama

seya-tu@pop.med.hokudai.ac.jp (TS); yamazas@med.nagoya-cu.ac.jp (SY)

Affiliations Department of Microbiology and Immunology, Graduate School of Medicine, Hokkaido University, Sapporo, Japan, Department of Geriatric and Environmental Dermatology, Nagoya City University, Graduate School of Medical Sciences, Nagoya, Japan

Akira Maruyama

Contributed equally to this work with: Sayuri Yamazaki, Akira Maruyama

Affiliation Department of Microbiology and Immunology, Graduate School of Medicine, Hokkaido University, Sapporo, Japan

Kohei Okada

Affiliation Department of Microbiology and Immunology, Graduate School of Medicine, Hokkaido University, Sapporo, Japan

Misako Matsumoto

Affiliation Department of Microbiology and Immunology, Graduate School of Medicine, Hokkaido University, Sapporo, Japan

Akimichi Morita

Affiliation Department of Geriatric and Environmental Dermatology, Nagoya City University, Graduate School of Medical Sciences, Nagoya, Japan

Tsukasa Seya

seya-tu@pop.med.hokudai.ac.jp (TS); yamazas@med.nagoya-cu.ac.jp (SY)

Affiliation Department of Microbiology and Immunology, Graduate School of Medicine, Hokkaido University, Sapporo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SY. Performed the experiments: A. Maruyama SY KO. Analyzed the data: SY A. Maruyama MM A. Morita TS. Wrote the paper: SY.