About the Authors

John E. Bowers

Affiliation Department of Plant Biology, University of Georgia, Athens, Georgia, United States of America

Savithri Nambeesan

Affiliation Department of Plant Biology, University of Georgia, Athens, Georgia, United States of America

Jonathan Corbi

Affiliation Department of Plant Biology, University of Georgia, Athens, Georgia, United States of America

Michael S. Barker

Affiliation Department of Ecology and Evolutionary Biology, University of Arizona, Tucson, Arizona, United States of America

Loren H. Rieseberg

Affiliations Department of Botany, University of British Columbia, Vancouver, British Columbia, Canada, Department of Biology, Indiana University, Bloomington, Indiana, United States of America

Steven J. Knapp

Current address: Monsanto Company, Woodland, California, United States of America

Affiliation Center for Applied Genetic Technologies, University of Georgia, Athens, Georgia, United States of America

John M. Burke

jmburke@uga.edu

Affiliation Department of Plant Biology, University of Georgia, Athens, Georgia, United States of America

Competing Interests

This study was supported in part by funding from commercial sources (Syngenta and Biogemma). This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SJK JEB LHR MSB JMB. Performed the experiments: JEB SN JC. Analyzed the data: JEB. Contributed reagents/materials/analysis tools: JMB. Wrote the paper: JEB JMB.