About the Authors

Marylene Y. Peroval

Affiliation Institute for Animal Health, Compton, Berkshire, United Kingdom

Amy C. Boyd

Affiliation The Jenner Institute, Old Road Campus Research Building, University of Oxford, Nuffield Department of Clinical Medicine, Oxford, Oxfordshire, United Kingdom

John R. Young

Affiliation Institute for Animal Health, Compton, Berkshire, United Kingdom

Adrian L. Smith

adrian.smith@zoo.ox.ac.uk

Affiliation Department of Zoology, University of Oxford, Tinbergen Building, Oxford, Oxfordshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Supervised all research: ALS. Conceived and designed the experiments: MYP ALS. Performed the experiments: MYP. Analyzed the data: MYP JRY ACB. Contributed reagents/materials/analysis tools: MYP JRY. Wrote the paper: ALS MYP ACB JRY.