About the Authors

Caitlyn W. Barrett

Affiliations Division of Gastroenterology, Department of Medicine, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Department of Cancer Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

J. Joshua Smith

Affiliation Department of Biostatistics, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Lauren C. Lu

Affiliation School for Science and Math, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Nicholas Markham

Affiliation Department of Cancer Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Kristy R. Stengel

Affiliation Department of Biochemistry, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Sarah P. Short

Affiliation Department of Cancer Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Baolin Zhang

Affiliations Division of Gastroenterology, Department of Medicine, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Department of Cancer Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Aubrey A. Hunt

Affiliation Department of Biochemistry, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Barbara M. Fingleton

Affiliations Department of Cancer Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Vanderbilt Ingram Cancer Center, Nashville, Tennessee, United States of America

Robert H. Carnahan

Affiliation Department of Cancer Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Michael E. Engel

Affiliation Huntsman Cancer Institute, University of Utah, Salt Lake City, United States of America

Xi Chen

Affiliations Department of Biostatistics, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Vanderbilt Ingram Cancer Center, Nashville, Tennessee, United States of America

R. Daniel Beauchamp

Affiliations Section of Surgical Sciences, Department of Surgery, Division of Surgical Oncology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Vanderbilt Ingram Cancer Center, Nashville, Tennessee, United States of America

Keith T. Wilson

Affiliations Division of Gastroenterology, Department of Medicine, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Department of Cancer Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Vanderbilt Ingram Cancer Center, Nashville, Tennessee, United States of America, Veterans Affairs Tennessee Valley Health Care System, Nashville, Tennessee, United States of America

Scott W. Hiebert

Affiliations Department of Biochemistry, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Vanderbilt Ingram Cancer Center, Nashville, Tennessee, United States of America

Albert B. Reynolds

Affiliations Department of Cancer Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Vanderbilt Ingram Cancer Center, Nashville, Tennessee, United States of America

Christopher S. Williams

christopher.williams@vanderbilt.edu

Affiliations Division of Gastroenterology, Department of Medicine, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Department of Cancer Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Vanderbilt Ingram Cancer Center, Nashville, Tennessee, United States of America, Veterans Affairs Tennessee Valley Health Care System, Nashville, Tennessee, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CWB CSW ABR SWH. Performed the experiments: CWB LCL NM KRS SPS. Analyzed the data: CWB JJS. Contributed reagents/materials/analysis tools: CWB JJS NM KRS BZ AAH BMF RHC MEE XC RDB. Wrote the paper: CWB JJS MEE KTW SWH ABR CSW.