About the Authors

James B. Stanton

jstanton@vetmed.wsu.edu

Affiliation Department of Veterinary Microbiology and Pathology, Washington State University, Pullman, Washington, United States of America

David A. Schneider

Affiliation United States Department of Agriculture, Agricultural Research Service, Animal Disease Research Unit, Pullman, Washington, United States of America

Kelcey D. Dinkel

Affiliation Department of Veterinary Microbiology and Pathology, Washington State University, Pullman, Washington, United States of America

Bethany F. Balmer

Affiliation Department of Veterinary Microbiology and Pathology, Washington State University, Pullman, Washington, United States of America

Timothy V. Baszler

Affiliation Department of Veterinary Microbiology and Pathology, Washington State University, Pullman, Washington, United States of America

Bruce A. Mathison

Affiliation Department of Veterinary Microbiology and Pathology, Washington State University, Pullman, Washington, United States of America

David W. Boykin

Affiliation Department of Chemistry, Georgia State University, Atlanta, Georgia, United States of America

Arvind Kumar

Affiliation Department of Chemistry, Georgia State University, Atlanta, Georgia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JBS DAS TVB DWB. Performed the experiments: JBS KDD BFB BAM AK. Analyzed the data: JBS DAS KDD BFB. Contributed reagents/materials/analysis tools: DWB AK. Wrote the paper: JBS DAS KDD TVB BAM DWB.