About the Authors

Andrew J. J. MacIntosh

macintosh@pri.kyoto-u.ac.jp

Affiliation Center for International Collaboration and Advanced Studies in Primatology, Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan

Armand Jacobs

Affiliation Département Ecologie, Physiologie et Ethologie, Institut Pluridisciplinaire Hubert Curien Centre National de la Recherche Scientifique UMR7178, Université de Strasbourg, Strasbourg, Alsace, France

Cécile Garcia

Affiliation Laboratoire de Dynamique de l’Évolution Humaine, Centre National de la Recherche Scientifique UPR 2147, Paris, Île-de-France, France

Keiko Shimizu

Affiliation Department of Zoology, Faculty of Science, Okayama University of Science, Okayama City, Okayama, Japan

Keiko Mouri

Affiliation Department of Zoology, Faculty of Science, Okayama University of Science, Okayama City, Okayama, Japan

Michael A. Huffman

Affiliation Center for International Collaboration and Advanced Studies in Primatology, Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan

Alexander D. Hernandez

Affiliation Center for Infectious Disease Dynamics, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AJJM ADH AJ. Performed the experiments: AJJM CG KM. Analyzed the data: AJJM AJ. Wrote the paper: AJJM ADH. Contributed substantially to supervision and discussion: MAH KS.