About the Authors

Wenbao Qi

Contributed equally to this work with: Wenbao Qi, Jin Tian

Affiliations College of Veterinary Medicine, South China Agricultural University, Guangdong, People's Republic of China, MOA Key Laboratory for Animal Vaccine Development, Guangzhou, People's Republic of China, Key Lab of Zoonoses Control and Prevention of Guangdong, Guangdong, People's Republic of China

Jin Tian

Contributed equally to this work with: Wenbao Qi, Jin Tian

Affiliations College of Veterinary Medicine, South China Agricultural University, Guangdong, People's Republic of China, MOA Key Laboratory for Animal Vaccine Development, Guangzhou, People's Republic of China, Key Lab of Zoonoses Control and Prevention of Guangdong, Guangdong, People's Republic of China

Changhui Zhang

Affiliations College of Veterinary Medicine, South China Agricultural University, Guangdong, People's Republic of China, MOA Key Laboratory for Animal Vaccine Development, Guangzhou, People's Republic of China, Key Lab of Zoonoses Control and Prevention of Guangdong, Guangdong, People's Republic of China

Jun He

Affiliations College of Veterinary Medicine, South China Agricultural University, Guangdong, People's Republic of China, MOA Key Laboratory for Animal Vaccine Development, Guangzhou, People's Republic of China, Key Lab of Zoonoses Control and Prevention of Guangdong, Guangdong, People's Republic of China

Zhangyong Ning

Affiliations College of Veterinary Medicine, South China Agricultural University, Guangdong, People's Republic of China, MOA Key Laboratory for Animal Vaccine Development, Guangzhou, People's Republic of China, Key Lab of Zoonoses Control and Prevention of Guangdong, Guangdong, People's Republic of China

Peirong Jiao

Affiliations College of Veterinary Medicine, South China Agricultural University, Guangdong, People's Republic of China, MOA Key Laboratory for Animal Vaccine Development, Guangzhou, People's Republic of China, Key Lab of Zoonoses Control and Prevention of Guangdong, Guangdong, People's Republic of China

Ming Liao

mliao@scau.edu.cn

Affiliations College of Veterinary Medicine, South China Agricultural University, Guangdong, People's Republic of China, MOA Key Laboratory for Animal Vaccine Development, Guangzhou, People's Republic of China, Key Lab of Zoonoses Control and Prevention of Guangdong, Guangdong, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JT WQ ML. Performed the experiments: JT JH. Analyzed the data: CZ JT. Contributed reagents/materials/analysis tools: ZN JT. Wrote the paper: JT PJ.