About the Authors

Christopher E. Heise

heise.christopher@gene.com (CEH); michelle.arkin@ucsf.edu (MRA)

Affiliation Department of Biochemical and Cellular Pharmacology, Genentech, Inc., South San Francisco, California, United States of America

Jeremy Murray

Affiliation Department of Structural Biology, Genentech, Inc., South San Francisco, California, United States of America

Katherine E. Augustyn

Affiliation Small Molecule Discovery Center, University of California San Francisco, San Francisco, California, United States of America

Brandon Bravo

Affiliation Department of Biochemical and Cellular Pharmacology, Genentech, Inc., South San Francisco, California, United States of America

Preeti Chugha

Affiliation Small Molecule Discovery Center, University of California San Francisco, San Francisco, California, United States of America

Frederick Cohen

Affiliation Department of Discovery Chemistry, Genentech, Inc., South San Francisco, California, United States of America

Anthony M. Giannetti

Affiliation Department of Biochemical and Cellular Pharmacology, Genentech, Inc., South San Francisco, California, United States of America

Paul Gibbons

Affiliation Department of Discovery Chemistry, Genentech, Inc., South San Francisco, California, United States of America

Rami N. Hannoush

Affiliation Department of Early Discovery Biochemistry, Genentech, Inc., South San Francisco, California, United States of America

Brian R. Hearn

Affiliation Small Molecule Discovery Center, University of California San Francisco, San Francisco, California, United States of America

Priyadarshini Jaishankar

Affiliation Small Molecule Discovery Center, University of California San Francisco, San Francisco, California, United States of America

Cuong Q. Ly

Affiliation Department of Discovery Chemistry, Genentech, Inc., South San Francisco, California, United States of America

Kinjalkumar Shah

Affiliation Department of Biochemical and Cellular Pharmacology, Genentech, Inc., South San Francisco, California, United States of America

Karen Stanger

Affiliation Department of Early Discovery Biochemistry, Genentech, Inc., South San Francisco, California, United States of America

Micah Steffek

Affiliation Department of Structural Biology, Genentech, Inc., South San Francisco, California, United States of America

Yinyan Tang

Affiliation Small Molecule Discovery Center, University of California San Francisco, San Francisco, California, United States of America

Xianrui Zhao

Affiliation Department of Discovery Chemistry, Genentech, Inc., South San Francisco, California, United States of America

Joseph W. Lewcock

Affiliation Department of Neuroscience, Genentech, Inc., South San Francisco, California, United States of America

Adam R. Renslo

Affiliation Small Molecule Discovery Center, University of California San Francisco, San Francisco, California, United States of America

John Flygare

Affiliation Department of Discovery Chemistry, Genentech, Inc., South San Francisco, California, United States of America

Michelle R. Arkin

heise.christopher@gene.com (CEH); michelle.arkin@ucsf.edu (MRA)

Affiliation Small Molecule Discovery Center, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

Several authors of this manuscript are employed by Genentech, Inc. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials. There are no patents, products in development, or marketed products to declare.

Author Contributions

Conceived and designed the experiments: CEH AMG RNH JM MS PG KEA FC BRH ARR MRA. Performed the experiments: CEH KEA BB AMG RNH K. Shah K. Stanger YT JM MS PG PC FC BRH PJ CQL XZ ARR. Analyzed the data: CEH KEA BB AMG RNH K. Shah K. Stanger YT JM PG MS FC BRH ARR MRA. Contributed reagents/materials/analysis tools: PG XZ CQL FC. Wrote the paper: CEH JM MRA FC ARR BRH AMG PG. Led the project team: JF JWL ARR MRA.