About the Authors

Paula Stewart

Affiliation Neurobiology Division, The Roslin Institute & R(D)SVS, University of Edinburgh, Roslin, United Kingdom

Lauren Campbell

Affiliation Neurobiology Division, The Roslin Institute & R(D)SVS, University of Edinburgh, Roslin, United Kingdom

Susan Skogtvedt

Affiliation Norwegian School of Veterinary Science, Dept. Basic Sciences & Aquatic Medicine, Oslo, Norway

Karen A. Griffin

Affiliation Wildlife Research Center, Colorado Division of Parks and Wildlife, Fort Collins, Colorado, United States of America

Jon M. Arnemo

Affiliation Department of Forestry & Wildlife Management, Faculty of Applied Ecology and Agricultural Sciences, Hedmark University College, Campus Evenstad, Elverum, Norway & Department of Wildlife, Fish and Environmental Studies, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Umeå, Sweden

Morten Tryland

Affiliations Norwegian School of Veterinary Science, Section of Arctic Veterinary Medicine, Tromsø, Norway, GenØk - Centre for Biosafety, Tromsø, Norway

Simon Girling

Affiliation Royal Zoological Society of Scotland, Edinburgh Zoo, Edinburgh, United Kingdom

Michael W. Miller

Affiliation Wildlife Research Center, Colorado Division of Parks and Wildlife, Fort Collins, Colorado, United States of America

Michael A. Tranulis

Affiliation Norwegian School of Veterinary Science, Dept. Basic Sciences & Aquatic Medicine, Oslo, Norway

Wilfred Goldmann

wilfred.goldmann@roslin.ed.ac.uk

Affiliation Neurobiology Division, The Roslin Institute & R(D)SVS, University of Edinburgh, Roslin, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PS MWM MAT WG. Performed the experiments: PS LC SS KAG JMA MT SG WG. Analyzed the data: PS LC SS MAT WG. Contributed reagents/materials/analysis tools: SS KAG JMA MT SG MWM MAT WG. Wrote the paper: PS MWM MAT.