About the Authors

Marion Favier

Affiliations University of Bordeaux, ISVV, Unit of Enology EA 4577, Villenave d'Ornon, France, SARCO, research subsidiary of the Laffort group, BP 40, Bordeaux, France

Eric Bilhère

Affiliation University of Bordeaux, ISVV, Unit of Enology EA 4577, Villenave d'Ornon, France

Aline Lonvaud-Funel

Affiliation University of Bordeaux, ISVV, Unit of Enology EA 4577, Villenave d'Ornon, France

Virginie Moine

Affiliation Laffort, BP 17, Bordeaux, France

Patrick M. Lucas

patrick.lucas@univ-bordeauxsegalen.fr

Affiliation University of Bordeaux, ISVV, Unit of Enology EA 4577, Villenave d'Ornon, France

Competing Interests

MF was a PhD student funded by SARCO, a subsidiary of the group Laffort. VM is an employee of the company Laffort. Consumables used in this work were for the largest part supported by funds from the University Bordeaux Segalen and a very minor part by Laffort. There are no patents, products in development or marketed products to declare. The authors declare no other competing interests. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MF EB PL. Performed the experiments: MF EB. Analyzed the data: MF EB AL VM PL. Contributed reagents/materials/analysis tools: AL VM PL. Wrote the paper: MF PL.