About the Authors

Laurel A. Kluber

laurel.kluber@ars.usda.gov

Affiliations Department of Biology, Case Western Reserve University, Cleveland, Ohio, United States of America, The Holden Arboretum, Kirtland, Ohio, United States of America

Sarah R. Carrino-Kyker

Affiliations Department of Biology, Case Western Reserve University, Cleveland, Ohio, United States of America, The Holden Arboretum, Kirtland, Ohio, United States of America

Kaitlin P. Coyle

Affiliation Department of Biology, Case Western Reserve University, Cleveland, Ohio, United States of America

Jared L. DeForest

Affiliation Department of Environmental and Plant Biology, Ohio University, Athens, Ohio, United States of America

Charlotte R. Hewins

Affiliation The Holden Arboretum, Kirtland, Ohio, United States of America

Alanna N. Shaw

Affiliation Department of Environmental and Plant Biology, Ohio University, Athens, Ohio, United States of America

Kurt A. Smemo

Affiliations The Holden Arboretum, Kirtland, Ohio, United States of America, Department of Biological Sciences, Kent State University, Kent, Ohio, United States of America

David J. Burke

Affiliations Department of Biology, Case Western Reserve University, Cleveland, Ohio, United States of America, The Holden Arboretum, Kirtland, Ohio, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JLD DJB KAS. Performed the experiments: LAK SRCK KPC JLD DJB KAS CRH ANS. Analyzed the data: LAK JLD SRCK. Wrote the paper: LAK SRCK KPC JLD DJB KAS.