About the Authors

Zi Long Wang

Contributed equally to this work with: Zi Long Wang, Ting Ting Liu

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, China

Ting Ting Liu

Contributed equally to this work with: Zi Long Wang, Ting Ting Liu

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, China

Zachary Y. Huang

Affiliations Department of Entomology, Michigan State University, East Lansing, Michigan, United States of America, Department of Ecology, Evolutionary Biology and Behavior Program, Michigan State University, East Lansing, Michigan, United States of America

Xiao Bo Wu

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, China

Wei Yu Yan

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, China

Zhi Jiang Zeng

bees1965@sina.com

Affiliation Honeybee Research Institute, Jiangxi Agricultural University, Nanchang, Jiangxi, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZJZ. Performed the experiments: TTL. Analyzed the data: ZLW. Contributed reagents/materials/analysis tools: WYY XBW. Wrote the paper: ZLW ZYH.