About the Authors

Gaia Nebbia

Affiliation Division of Infection and Immunity, University College London, London, United Kingdom

Dimitra Peppa

Affiliation Division of Infection and Immunity, University College London, London, United Kingdom

Anna Schurich

Affiliation Division of Infection and Immunity, University College London, London, United Kingdom

Pooja Khanna

Affiliations Division of Infection and Immunity, University College London, London, United Kingdom, Centre for Hepatology, University College London, London, United Kingdom

Harsimran D. Singh

Affiliation Centre for Hepatology, University College London, London, United Kingdom

Yang Cheng

Affiliation Division of Infection and Immunity, University College London, London, United Kingdom

William Rosenberg

Affiliation Centre for Hepatology, University College London, London, United Kingdom

Geoffrey Dusheiko

Affiliation Centre for Hepatology, University College London, London, United Kingdom

Richard Gilson

Affiliation Centre for Sexual Health and HIV, University College London, London, United Kingdom

Joanne ChinAleong

Affiliation Centre for Digestive Diseases, Institute of Cell and Molecular Science, Barts and the London School of Medicine and Dentistry, London, United Kingdom

Patrick Kennedy

Affiliation Centre for Digestive Diseases, Institute of Cell and Molecular Science, Barts and the London School of Medicine and Dentistry, London, United Kingdom

Mala K. Maini

m.maini@ucl.ac.uk

Affiliation Division of Infection and Immunity, University College London, London, United Kingdom

Competing Interests

This work was partially funded by an unrestricted the authors’ adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Critical review of manuscript: AS PK HDS WR GD RJG JC PTFK. Conceived and designed the experiments: GN DP MKM. Performed the experiments: GN DP AS HDS PK YC. Analyzed the data: GN DP AS HDS JC YC MKM. Contributed reagents/materials/analysis tools: PK WR GD RJG PTFK. Wrote the paper: GN DP MKM.