About the Authors

Jeffrey A. Butler

Contributed equally to this work with: Jeffrey A. Butler, Robert J. Mishur

Affiliations The Barshop Institute for Longevity and Aging Studies, University of Texas Health Science Center at San Antonio, San Antonio, Texas, United States of America, Department of Physiology, University of Texas Health Science Center at San Antonio, San Antonio, Texas, United States of America

Robert J. Mishur

Contributed equally to this work with: Jeffrey A. Butler, Robert J. Mishur

Affiliations The Barshop Institute for Longevity and Aging Studies, University of Texas Health Science Center at San Antonio, San Antonio, Texas, United States of America, Department of Physiology, University of Texas Health Science Center at San Antonio, San Antonio, Texas, United States of America

Alex F. Bokov

Affiliation The Barshop Institute for Longevity and Aging Studies, University of Texas Health Science Center at San Antonio, San Antonio, Texas, United States of America

Kevin W. Hakala

Affiliation Department of Biochemistry, University of Texas Health Science Center at San Antonio, San Antonio, Texas, United States of America

Susan T. Weintraub

Affiliation Department of Biochemistry, University of Texas Health Science Center at San Antonio, San Antonio, Texas, United States of America

Shane L. Rea

reas3@uthscsa.edu

Affiliations The Barshop Institute for Longevity and Aging Studies, University of Texas Health Science Center at San Antonio, San Antonio, Texas, United States of America, Department of Physiology, University of Texas Health Science Center at San Antonio, San Antonio, Texas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SLR JAB STW. Performed the experiments: JAB RJM KWH SLR. Analyzed the data: JAB RJM AFB SLR. Wrote the paper: RJM JAB STW SLR.