About the Authors

Robert A. Nawrocki

Affiliation Department of Computer Engineering, University of Denver, Denver, Colorado, United States of America

Majid Shaalan

Affiliation Department of Computer Science and Engineering, University of Colorado Denver, Denver, Colorado, United States of America

Sean E. Shaheen

Affiliation Department of Physics and Astronomy, University of Denver, Denver, Colorado, United States of America

Nancy M. Lorenzon

nancy.lorenzon@du.edu

Affiliation Department of Biological Sciences, University of Denver, Denver, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RAN MS SES NML. Performed the experiments: RAN MS NML SES. Analyzed the data: RAN MS NML SES. Contributed reagents/materials/analysis tools: RAN MS NML SES. Wrote the paper: RAN MS NML SES. Inspiration for the study: RAN. Design of the cerebellar microzone model implemented in EmergentTM: RAN MS NML. Design of the puff & buzz conditioning training: RAN SES NML. Simulation of the damage experiment: RAN MS. Data analysis: RAN MS SES NML. Drafting the entirety of the document/manuscript: RAN. Drafting sections of the document/manuscript: MS. Correction of sections of the document/manuscript: SES. Drafting and correction of sections of the document/manuscript: NML.