About the Authors

Fernando A. Villanea

fervillanea@wsu.edu

Affiliations School of Biological Sciences, Washington State University, Pullman, Washington, United States of America, Department of Anthropology, University of California Santa Cruz, Santa Cruz, California, United States of America, Escuela de Biología, Universidad de Costa Rica, San José, Costa Rica

George H. Perry

Affiliation Department of Anthropology, Pennsylvania State University, University Park, Pennsylvania, United States of America

Gustavo A. Gutiérrez-Espeleta

Affiliation Escuela de Biología, Universidad de Costa Rica, San José, Costa Rica

Nathaniel J. Dominy

Affiliations Department of Anthropology, Dartmouth College, Hanover, New Hampshire, United States of America, Department of Biological Sciences, Dartmouth College, Hanover, New Hampshire, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FAV NJD GHP. Performed the experiments: FAV. Analyzed the data: FAV. Contributed reagents/materials/analysis tools: GG. Wrote the paper: FAV NJD GHP.