About the Authors

W. David Walter

wdwalter@psu.edu

Current address: US Geological Survey, Pennsylvania Cooperative Fish and Wildlife Research Unit, University Park, Pennsylvania, United States of America

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Services, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Charles W. Anderson

Current address: Missouri Department of Conservation, West Plains, Missouri, United States of America

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Services, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Kurt C. VerCauteren

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Services, Wildlife Services, National Wildlife Research Center, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WDW CWA KCV. Performed the experiments: WDW CWA. Analyzed the data: WDW. Contributed reagents/materials/analysis tools: WDW CWA KCV. Wrote the paper: WDW CWA KCV.