About the Authors

Tomoaki Koga

Contributed equally to this work with: Tomoaki Koga, Yukari Kai

Affiliations Department of Molecular Medicine, Graduate School of Pharmaceutical Sciences and Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan, Research Fellow of the Japan Society for the Promotion of Science (JSPS), Tokyo, Japan, Department of Medical Biochemistry, Graduate School of Medical Sciences, Kyushu University, Fukuoka, Japan

Yukari Kai

Contributed equally to this work with: Tomoaki Koga, Yukari Kai

Affiliation Department of Molecular Medicine, Graduate School of Pharmaceutical Sciences and Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan

Ryosuke Fukuda

Affiliation Department of Molecular Medicine, Graduate School of Pharmaceutical Sciences and Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan

Saori Morino-Koga

Affiliation Department of Molecular Medicine, Graduate School of Pharmaceutical Sciences and Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan

Mary Ann Suico

Affiliation Department of Molecular Medicine, Graduate School of Pharmaceutical Sciences and Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan

Kosuke Koyama

Affiliation Department of Molecular Medicine, Graduate School of Pharmaceutical Sciences and Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan

Takashi Sato

Affiliation Department of Molecular Medicine, Graduate School of Pharmaceutical Sciences and Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan

Tsuyoshi Shuto

Affiliation Department of Molecular Medicine, Graduate School of Pharmaceutical Sciences and Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan

Hirofumi Kai

hirokai@gpo.kumamoto-u.ac.jp

Affiliation Department of Molecular Medicine, Graduate School of Pharmaceutical Sciences and Global COE “Cell Fate Regulation Research and Education Unit”, Kumamoto University, Kumamoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TK HK. Performed the experiments: TK YK RF KK. Analyzed the data: TK SMK T. Sato HK. Contributed reagents/materials/analysis tools: T. Shuto HK. Wrote the paper: TK MAS HK.