About the Authors

Shing Hwa Liu

Contributed equally to this work with: Shing Hwa Liu, Chin Chang Shen

Affiliation Institute of Toxicology, College of Medicine, National Taiwan University, Taipei, Taiwan

Keh Bin Wang

Affiliations Department of Nuclear Medicine, Kuang Tien General Hospital, Taichung, Taiwan, Institute of Biomedical Sciences, National Chung Hsing University, Taichung, Taiwan

Keng Hsin Lan

Affiliation Division of Gastroenterology, Department of Medicine, Taipei Veterans General Hospital, Taipei, Taiwan

Wen Jane Lee

Affiliation Department of Education and Research, Taichung Veterans General Hospital, Taichung, Taiwan

Hung Chuan Pan

Affiliations Department of Education and Research, Taichung Veterans General Hospital, Taichung, Taiwan, Institute of Biomedical Sciences, National Chung Hsing University, Taichung, Taiwan

Sheng Mao Wu

Affiliation Institute of Biomedical Sciences, National Chung Hsing University, Taichung, Taiwan

Yen Chun Peng

Affiliation Division of Gastroenterology, Department of Internal Medicine, Taichung Veterans General Hospital, Taichung, Taiwan

Yi Ching Chen

Affiliation Institute of Biomedical Sciences, National Chung Hsing University, Taichung, Taiwan

Chin Chang Shen

Contributed equally to this work with: Shing Hwa Liu, Chin Chang Shen

Affiliations Department of Chemistry, National Taiwan University, Taipei, Taiwan, Institute of Nuclear Energy Research, Atomic Energy Council, Taoyuan, Taiwan

Hsu Chen Cheng

Affiliation Department of life Sciences, National Chung Hsing University, Taichung, Taiwan

Ko Kaung Liao

Affiliation Department of Anatomy, Chung Shan Medical University, Taichung, Taiwan

Meei Ling Sheu

mlsheu@nchu.edu.tw

Affiliations Department of Education and Research, Taichung Veterans General Hospital, Taichung, Taiwan, Institute of Biomedical Sciences, National Chung Hsing University, Taichung, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SHL MLS. Performed the experiments: KBW KHL CCS SMW YCP YCC KKL. Analyzed the data: SHL MLS. Contributed reagents/materials/analysis tools: KKL CCS WJL HCP HCC. Wrote the paper: SHL MLS.