About the Authors

Han-Lin Huang

Contributed equally to this work with: Han-Lin Huang, Hsueh-Yun Lee

Affiliation Pharmacological Institute, College of Medicine, National Taiwan University, Taipei, Taiwan

Hsueh-Yun Lee

Contributed equally to this work with: Han-Lin Huang, Hsueh-Yun Lee

Affiliation School of Pharmacy, College of Pharmacy, Taipei Medical University, Taipei, Taiwan

An-Chi Tsai

Affiliation Institute of Biotechnology and Pharmaceutical Research, National Health Research Institutes, Zhunan Town, Miaoli County, Taiwan

Chieh-Yu Peng

Affiliation School of Chinese Medicine, College of Chinese Medicine, China Medical University, Taichung, Taiwan

Mei-Jung Lai

Affiliation School of Pharmacy, College of Pharmacy, Taipei Medical University, Taipei, Taiwan

Jing-Chi Wang

Affiliation Institute of Biotechnology and Pharmaceutical Research, National Health Research Institutes, Zhunan Town, Miaoli County, Taiwan

Shiow-Lin Pan

jpl@tmu.edu.tw (JPL); apan0826@nhri.org.tw (SLP); cmteng@ntu.edu.tw (CMT)

Affiliations Institute of Biotechnology and Pharmaceutical Research, National Health Research Institutes, Zhunan Town, Miaoli County, Taiwan, Department of Pharmacology, Taipei Medical University, Taipei, Taiwan

Che-Ming Teng

jpl@tmu.edu.tw (JPL); apan0826@nhri.org.tw (SLP); cmteng@ntu.edu.tw (CMT)

Affiliation Pharmacological Institute, College of Medicine, National Taiwan University, Taipei, Taiwan

Jing-Ping Liou

jpl@tmu.edu.tw (JPL); apan0826@nhri.org.tw (SLP); cmteng@ntu.edu.tw (CMT)

Affiliation School of Pharmacy, College of Pharmacy, Taipei Medical University, Taipei, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HLH HYL SLP CMT JPL. Performed the experiments: HLH HYL SLP CMT JPL. Analyzed the data: HLH HYL ACT CYP MJL JCW. Contributed reagents/materials/analysis tools: HLH HYL ACT CYP MJL JCW SLP JPL. Wrote the paper: HLH SLP CMT JPL.