About the Authors

Helga H. Olsen

helga.haugomolsen@mpa.se

Affiliation Department of Medicine, Division of Respiratory Medicine, Karolinska University Hospital, Solna, Stockholm, Sweden

Johan Grunewald
Göran Tornling
C. Magnus Sköld
Anders Eklund

Competing Interests

Funding was received from Aztra Zeneca. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HO JG GT CMS AE. Performed the experiments: CMS AE. Analyzed the data: HO JG GT CMS AE. Wrote the paper: HO. Critically revising the article: JG GT CMS AE.