About the Authors

Ripla Arora

Affiliation Department of Genetics and Development, Columbia University Medical Center, New York, New York, United States of America

Chelsea M. del Alcazar

Affiliation Department of Genetics and Development, Columbia University Medical Center, New York, New York, United States of America

Edward E. Morrisey

Affiliation Department of Medicine and Cell and Developmental Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

L. A. Naiche

Affiliation Cancer and Developmental Biology Laboratory, National Cancer Institute, Frederick, Maryland, United States of America

Virginia E. Papaioannou

vep1@columbia.edu

Affiliation Department of Genetics and Development, Columbia University Medical Center, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RA VEP. Performed the experiments: RA CMdA LAN. Analyzed the data: RA CMdA VEP. Contributed reagents/materials/analysis tools: EEM VEP. Wrote the paper: RA VEP.