About the Authors

Wei Zhu

Contributed equally to this work with: Wei Zhu, Qing Miao, Dan Sun

Affiliation State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian, Shandong, People’s Republic of China

Qing Miao

Contributed equally to this work with: Wei Zhu, Qing Miao, Dan Sun

Affiliation State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian, Shandong, People’s Republic of China

Dan Sun

Contributed equally to this work with: Wei Zhu, Qing Miao, Dan Sun

Affiliation State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian, Shandong, People’s Republic of China

Guodong Yang

Affiliation State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian, Shandong, People’s Republic of China

Changai Wu

Affiliation State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian, Shandong, People’s Republic of China

Jinguang Huang

Affiliation State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian, Shandong, People’s Republic of China

Chengchao Zheng

cczheng@sdau.edu.cn

Affiliation State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian, Shandong, People’s Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WZ GY CW JH CZ. Performed the experiments: WZ QM DS. Analyzed the data: WZ QM DS. Contributed reagents/materials/analysis tools: CW JH CZ. Wrote the paper: WZ QM DS.