About the Authors

Etsuro Ito

eito@kph.bunri-u.ac.jp

Affiliation Laboratory of Functional Biology, Kagawa School of Pharmaceutical Sciences, Tokushima Bunri University, Sanuki, Japan

Emi Otsuka

Affiliation Laboratory of Functional Biology, Kagawa School of Pharmaceutical Sciences, Tokushima Bunri University, Sanuki, Japan

Noriyuki Hama

Current address: Noriyuki Hama, Department of Neural and Muscular Physiology, Shimane University School of Medicine, Izumo, Japan

Affiliation Division of Biological Sciences, Graduate School of Science, Hokkaido University, Sapporo, Japan

Hitoshi Aonuma

Affiliation Complex Systems Research Group, Research Institute for Electronic Science, Hokkaido University, Sapporo, Japan

Ryuichi Okada

Affiliation Laboratory of Functional Biology, Kagawa School of Pharmaceutical Sciences, Tokushima Bunri University, Sanuki, Japan

Dai Hatakeyama

Affiliation Laboratory of Functional Biology, Kagawa School of Pharmaceutical Sciences, Tokushima Bunri University, Sanuki, Japan

Yutaka Fujito

Affiliation Department of Systems Neuroscience, School of Medicine, Sapporo Medical University, Sapporo, Japan

Suguru Kobayashi

Affiliation Laboratory of Functional Biology, Kagawa School of Pharmaceutical Sciences, Tokushima Bunri University, Sanuki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EI. Performed the experiments: EI EO NH HA. Analyzed the data: EI EO RO DH SK. Wrote the paper: EI EO NH HA RO DH YF SK.