About the Authors

Tomoyasu Sugiyama

Contributed equally to this work with: Tomoyasu Sugiyama, Rie Sugioka-Sugiyama

sugiyamt@biol.tsukuba.ac.jp

Affiliations Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan, Initiative for the Promotion of Young Scientists' Independent Research, University of Tsukuba, Tsukuba, Ibaraki, Japan, Precursory Research for Embryonic Science and Technology (PRESTO), Japan Science and Technology Agency (JST), Kawaguchi, Saitama, Japan

Rie Sugioka-Sugiyama

Contributed equally to this work with: Tomoyasu Sugiyama, Rie Sugioka-Sugiyama

Affiliations Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan, Initiative for the Promotion of Young Scientists' Independent Research, University of Tsukuba, Tsukuba, Ibaraki, Japan

Kazumasa Hada

Affiliations Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan, Initiative for the Promotion of Young Scientists' Independent Research, University of Tsukuba, Tsukuba, Ibaraki, Japan

Ryusuke Niwa

Affiliations Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan, Initiative for the Promotion of Young Scientists' Independent Research, University of Tsukuba, Tsukuba, Ibaraki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TS RS-S. Performed the experiments: TS RS-S KH RN. Analyzed the data: TS RS-S KH RN. Contributed reagents/materials/analysis tools: TS RS-S KH RN. Wrote the paper: TS RS-S RN.