About the Authors

Chin-Yu Lai

Affiliation Pharmacological Institute, College of Medicine, National Taiwan University, Taipei, Taiwan

An-Chi Tsai

Affiliation Institute of Biotechnology and Pharmaceutical Research, National Health Research Institutes, Zhunan Town, Taiwan

Mei-Chuan Chen

Affiliation Pharmacological Institute, College of Medicine, National Taiwan University, Taipei, Taiwan

Li-Hsun Chang

Affiliation Pharmacological Institute, College of Medicine, National Taiwan University, Taipei, Taiwan

Hui-Lung Sun

Affiliation Pharmacological Institute, College of Medicine, National Taiwan University, Taipei, Taiwan

Ya-Ling Chang

Affiliation Pharmacological Institute, College of Medicine, National Taiwan University, Taipei, Taiwan

Chien-Chih Chen

Affiliation Department of Biotechnology, Hungkuang University, Taichung, Taiwan

Che-Ming Teng

Contributed equally to this work with: Che-Ming Teng, Shiow-Lin Pan

cmteng@ntu.edu.tw (CMT); apan0826@nhri.org.tw (SLP)

Affiliation Pharmacological Institute, College of Medicine, National Taiwan University, Taipei, Taiwan

Shiow-Lin Pan

Contributed equally to this work with: Che-Ming Teng, Shiow-Lin Pan

cmteng@ntu.edu.tw (CMT); apan0826@nhri.org.tw (SLP)

Affiliation Institute of Biotechnology and Pharmaceutical Research, National Health Research Institutes, Zhunan Town, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CYL SLP CMT ACT MCC. Performed the experiments: CYL ACT MCC. Analyzed the data: CYL SLP CMT LHC HLS. Contributed reagents/materials/analysis tools: CCC YLC LHC HLS CYL. Wrote the paper: SLP CMT CYL.