About the Authors

Jung Eun Park

Affiliation Department of Theriogenology & Biotechnology, College of Veterinary Medicine, Seoul National University, Seoul, Republic of Korea

Min Jung Kim

Affiliation Department of Theriogenology & Biotechnology, College of Veterinary Medicine, Seoul National University, Seoul, Republic of Korea

Seung Kwon Ha

Affiliation Life Science of R&D Center, SK chemicals, Bundang-gu, Seongnam-si, Gyeonggi-do, Seoul, Republic of Korea

So Gun Hong

Affiliation Department of Theriogenology & Biotechnology, College of Veterinary Medicine, Seoul National University, Seoul, Republic of Korea

Hyun Ju Oh

Affiliation Department of Theriogenology & Biotechnology, College of Veterinary Medicine, Seoul National University, Seoul, Republic of Korea

Geon A. Kim

Affiliation Department of Theriogenology & Biotechnology, College of Veterinary Medicine, Seoul National University, Seoul, Republic of Korea

Eun Jung Park

Affiliation Department of Theriogenology & Biotechnology, College of Veterinary Medicine, Seoul National University, Seoul, Republic of Korea

Jung Taek Kang

Affiliation Department of Theriogenology & Biotechnology, College of Veterinary Medicine, Seoul National University, Seoul, Republic of Korea

Islam M. Saadeldin

Affiliations Department of Theriogenology & Biotechnology, College of Veterinary Medicine, Seoul National University, Seoul, Republic of Korea, Department of Physiology, Faculty of Veterinary Medicine, Zagazig University, Zagazig, Sharkia, Egypt

Goo Jang

Affiliation Department of Theriogenology & Biotechnology, College of Veterinary Medicine, Seoul National University, Seoul, Republic of Korea

Byeong Chun Lee

bclee@snu.ac.kr

Affiliation Department of Theriogenology & Biotechnology, College of Veterinary Medicine, Seoul National University, Seoul, Republic of Korea

Competing Interests

Nestlé Purina PetCare provided financial support, and Seung Kwon Ha is an employee of ORIENT CRO. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: JEP MJK SKH IMS BCL. Performed the experiments: JEP MJK SKH SGH HJO GAK EJP JTK GJ BCL. Analyzed the data: JEP SKH. Contributed reagents/materials/analysis tools: JEP SKH. Wrote the paper: JEP SKH.