About the Authors

Heinz Ruffner

heinz.ruffner@novartis.com (HR); feng.cong@novartis.com (FC)

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Joëlle Sprunger

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Olga Charlat

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Juliet Leighton-Davies

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Bianka Grosshans

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Adrian Salathe

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Svenja Zietzling

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Valérie Beck

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Maxime Therier

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Andrea Isken

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Yang Xie

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Yue Zhang

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Huaixiang Hao

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Xiaoying Shi

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Dong Liu

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Qinhui Song

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Ieuan Clay

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Gabriele Hintzen

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Jan Tchorz

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Laure C. Bouchez

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Gregory Michaud

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Peter Finan

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Vic E. Myer

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Tewis Bouwmeester

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Jeff Porter

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Marc Hild

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Fred Bassilana

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Christian N. Parker

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Novartis Pharma AG, Postfach, Basel, Switzerland

Feng Cong

heinz.ruffner@novartis.com (HR); feng.cong@novartis.com (FC)

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Competing Interests

The authors are employed by Novartis Institutes for Biomedical Research. This does not alter the authors’ adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HR GH JT LCB GM PF DL VEM TB JP MH FB CNP FC. Performed the experiments: JS OC JL BG AS SZ VB MT AI YX YZ HH XS QS IC. Analyzed the data: HR MHFB CNP FC. Wrote the paper: HR FB CNP FC.