About the Authors

Neil Beeharry

Neil.Beeharry@fccc.edu (NB); Timothy.Yen@fccc.edu (TJY)

Affiliation Basic Science Division, Fox Chase Cancer Center, Philadelphia, Pennsylvania, United States of America

Jerome B. Rattner

Affiliation Department of Cell Biology and Anatomy, University of Calgary, Calgary, Alberta, Canada

Alfonso Bellacosa

Affiliation Basic Science Division, Fox Chase Cancer Center, Philadelphia, Pennsylvania, United States of America

Mitchell R. Smith

Affiliation Basic Science Division, Fox Chase Cancer Center, Philadelphia, Pennsylvania, United States of America

Timothy J. Yen

Neil.Beeharry@fccc.edu (NB); Timothy.Yen@fccc.edu (TJY)

Affiliation Basic Science Division, Fox Chase Cancer Center, Philadelphia, Pennsylvania, United States of America

Competing Interests

M.R. Smith is on the advisory board for Cephalon. No financial support was provided from Cephalon. There were no products/patents from Cephalon for these studies. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: NB MRS TJY. Performed the experiments: NB JBR TJY. Analyzed the data: NB TJY. Contributed reagents/materials/analysis tools: NB AB JBR MRS TJY. Wrote the paper: NB TJY.