About the Authors

Toritse Orubu

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Naif Khalaf Alharbi

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Teresa Lambe

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Sarah C. Gilbert

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Matthew G. Cottingham

matt.cottingham@ndm.ox.ac.uk

Affiliation The Jenner Institute, University of Oxford, Oxford, United Kingdom

Competing Interests

MGC is a named inventor on a patent application relating to the method described (United Kingdom Patent Application No. 1006405.3 “Poxvirus expression system”). SCG is a named inventor on a patent relating to prime-boost vaccination (WO9856919 “Methods and reagents for vaccination which generate a CD8 T cell immune response”). This does not alter the authors’ adherence to all the PLoS ONE policies on sharing data and materials..

Author Contributions

Conceived and designed the experiments: TO TL MGC. Performed the experiments: TO NKA. Analyzed the data: TO TL MGC . Wrote the paper: TO MGC. Discussed the experiments and reviewed and contributed to the manuscript: SCG.