About the Authors

Xin Huang

Contributed equally to this work with: Xin Huang, Kushi Kushekhar

Affiliations Department of Pathology and Medical Biology, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands, Department of Pathology, Health Science Center, Peking University, Beijing, China

Kushi Kushekhar

Contributed equally to this work with: Xin Huang, Kushi Kushekhar

Affiliation Department of Pathology and Medical Biology, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Ilja Nolte

Affiliation Department of Epidemiology, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Wierd Kooistra

Affiliation Department of Pathology and Medical Biology, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Lydia Visser

Affiliation Department of Pathology and Medical Biology, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Ilby Bouwman

Affiliation Department of Laboratory Medicine, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Niels Kouprie

Affiliation Department of Laboratory Medicine, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Rianne Veenstra

Affiliation Department of Pathology and Medical Biology, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Gustaaf van Imhoff

Affiliation Department of Hematology, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Bianca Olver

Affiliation Section of Cancer Genetics, Institute of Cancer Research, Sutton, United Kingdom

Richard S. Houlston

Affiliation Section of Cancer Genetics, Institute of Cancer Research, Sutton, United Kingdom

Sibrand Poppema

Affiliation Department of Pathology and Medical Biology, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Arjan Diepstra

Affiliation Department of Pathology and Medical Biology, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Bouke Hepkema

Affiliation Department of Laboratory Medicine, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Anke van den Berg

a.van.den.berg01@umcg.nl

Affiliation Department of Pathology and Medical Biology, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XH AB BH AD LV SP. Performed the experiments: XH WK RV NK LV IB. Analyzed the data: IN KK. Contributed reagents/materials/analysis tools: RSH BO IN SP GI. Wrote the paper: BH IN AD KK AB GI. Reclassification of patients: AD SP.