About the Authors

Sarah E. Pabian

sarah.pabian@colostate.edu

a Current address: Biology Department, Colorado State University, Fort Collins, Colorado, United States of America

Affiliation School of Forest Resources, Pennsylvania State University, University Park, Pennsylvania, United States of America

Nathan M. Ermer

b Current address: New York State Department of Environmental Conservation, Division of Fish, Wildlife, and Marine Resources, New Paltz, New York, United States of America

Affiliation School of Forest Resources, Pennsylvania State University, University Park, Pennsylvania, United States of America

Walter M. Tzilkowski

Affiliation School of Forest Resources, Pennsylvania State University, University Park, Pennsylvania, United States of America

Margaret C. Brittingham

Affiliation School of Forest Resources, Pennsylvania State University, University Park, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SEP NME WMT MCB. Performed the experiments: SEP NME. Analyzed the data: SEP NME WMT MCB. Wrote the paper: SEP NME WMT MCB.