About the Authors

Nicola M. Heller

Affiliation Department of Microbiology and Immunology and the Center for Vascular and Inflammatory Diseases, The University of Maryland School of Medicine, Baltimore, Maryland, United States of America

William M. Gwinn

Affiliation Department of Microbiology, Immunology, and Tropical Medicine, The George Washington University Medical Center, Washington, District of Columbia, United States of America

Raymond P. Donnelly

Affiliation Division of Therapeutic Proteins, Center for Drug Evaluation and Research, U.S. Food and Drug Administration, Bethesda, Maryland, United States of America

Stephanie L. Constant

Affiliation Department of Microbiology, Immunology, and Tropical Medicine, The George Washington University Medical Center, Washington, District of Columbia, United States of America

Achsah D. Keegan

akeegan@som.umaryland.edu

Affiliation Department of Microbiology and Immunology and the Center for Vascular and Inflammatory Diseases, The University of Maryland School of Medicine, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ADK SLC NMH. Performed the experiments: NMH WMG. Analyzed the data: NMH ADK RPD. Contributed reagents/materials/analysis tools: RPD WMG SLC. Wrote the paper: NMH ADK.