About the Authors

Holly J. Puglis

hpuglis@usgs.gov

Affiliation Department of Zoology, Miami University, Oxford, Ohio, United States of America

Michelle D. Boone

Affiliation Department of Zoology, Miami University, Oxford, Ohio, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HJP MDB. Performed the experiments: HJP. Analyzed the data: HJP MDB. Contributed reagents/materials/analysis tools: HJP MDB. Wrote the paper: HJP MDB.