About the Authors

Ken-ichi Inoue

Contributed equally to this work with: Ken-ichi Inoue, Daisuke Koketsu

Affiliation Systems Neuroscience Section, Primate Research Institute, Kyoto University, Inuyama, Japan

Daisuke Koketsu

Contributed equally to this work with: Ken-ichi Inoue, Daisuke Koketsu

Affiliation Division of System Neurophysiology, National Institute for Physiological Sciences and Department of Physiological Sciences, The Graduate University for Advanced Studies, Okazaki, Japan

Shigeki Kato

Affiliation Department of Molecular Genetics, Institute of Biomedical Sciences, Fukushima Medical University School of Medicine, Fukushima, Japan

Kazuto Kobayashi

Affiliation Department of Molecular Genetics, Institute of Biomedical Sciences, Fukushima Medical University School of Medicine, Fukushima, Japan

Atsushi Nambu

nambu@nips.ac.jp (AN); takada@pri.kyoto-u.ac.jp (MT)

Affiliation Division of System Neurophysiology, National Institute for Physiological Sciences and Department of Physiological Sciences, The Graduate University for Advanced Studies, Okazaki, Japan

Masahiko Takada

nambu@nips.ac.jp (AN); takada@pri.kyoto-u.ac.jp (MT)

Affiliation Systems Neuroscience Section, Primate Research Institute, Kyoto University, Inuyama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MT AN KI DK. Performed the experiments: KI DK AN. Analyzed the data: DK KI AN. Contributed reagents/materials/analysis tools: KK SK. Wrote the paper: MT AN KI DK KK.