About the Authors

Jesús Osorio

Affiliation Animal Health Department, Faculty of Veterinary Science, University of León, León, Spain

Ana Carvajal

Affiliation Animal Health Department, Faculty of Veterinary Science, University of León, León, Spain

Germán Naharro

Affiliation Animal Health Department, Faculty of Veterinary Science, University of León, León, Spain

Tom La

Affiliation School of Veterinary and Biomedical Science, Murdoch University, Murdoch, Western Australia, Australia

Nyree D. Phillips

Affiliation School of Veterinary and Biomedical Science, Murdoch University, Murdoch, Western Australia, Australia

Pedro Rubio

Affiliation Animal Health Department, Faculty of Veterinary Science, University of León, León, Spain

David J. Hampson

d.hampson@murdoch.edu.au

Affiliation School of Veterinary and Biomedical Science, Murdoch University, Murdoch, Western Australia, Australia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JO AC GN PR DJH. Performed the experiments: JO TL NDP. Analyzed the data: JO AC TL DJH. Contributed reagents/materials/analysis tools: JO AC GN NDP PR. Wrote the paper: JO AC DJH.