About the Authors

William H. Broach

Affiliation Department of Biological Sciences, Molecular and Cellular Biology Program, Ohio University, Athens, Ohio, United States of America

Nicholas Egan

Affiliation School of Life Sciences, University of Nevada, Las Vegas, Las Vegas, Nevada, United States of America

Helen J. Wing

Affiliation School of Life Sciences, University of Nevada, Las Vegas, Las Vegas, Nevada, United States of America

Shelley M. Payne

Affiliation Section of Molecular Genetics and Microbiology, The University of Texas at Austin, Austin, Texas, United States of America

Erin R. Murphy

murphye@ohio.edu

Affiliation Department of Biomedical Sciences, Ohio University Heritage College of Osteopathic Medicine, Athens, Ohio, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WHB HJW SMP ERM. Performed the experiments: WHB NE ERM. Analyzed the data: WHB NE HJW SMP ERM. Contributed reagents/materials/analysis tools: HJW SMP ERM. Wrote the paper: WHB HJW SMP ERM.