About the Authors

Takahisa Kawaguchi

Affiliations Center for Genomic Medicine, Kyoto University Graduate School of Medicine, Kyoto, Japan, Institut National de la Sante et de la Recherche Medicale (INSERM) Unite U852, Kyoto University Graduate School of Medicine, Kyoto, Japan

Yoshio Sumida

Affiliation Center for Digestive and Liver Diseases, Nara City Hospital, Nara, Japan

Atsushi Umemura

Affiliation Center of Gastroenterology and Hepatology, Saiseikai Suita Hospital, Suita, Japan

Keitaro Matsuo

Affiliation Division of Epidemiology and Prevention, Aichi Cancer Center, Nagoya, Japan

Meiko Takahashi

Affiliation Center for Genomic Medicine, Kyoto University Graduate School of Medicine, Kyoto, Japan

Toshinari Takamura

Affiliation Department of Disease Control and Homeostasis, Kanazawa University, Graduate School of Medical Science, Kanazawa, Japan

Kohichiroh Yasui

Affiliation Department of Molecular Gastroenterology and Hepatology, Graduate School of Medical Science, Kyoto Prefectural University of Medicine, Kyoto, Japan

Toshiji Saibara

Affiliation Department of Gastroenterology and Hepatology, Kochi Medical School, Kochi, Japan

Etsuko Hashimoto

Affiliation Department of Internal Medicine and Gastroenterology, Tokyo Women’s Medical University, Tokyo, Japan

Miwa Kawanaka

Affiliation Center of Liver Diseases, Kawasaki Hospital, Kawasaki Medical School, Okayama, Japan

Sumio Watanabe

Affiliation Department of Gastroenterology, Juntendo University School of Medicine, Tokyo, Japan

Sumio Kawata

Affiliation Department of Gastroenterology, Yamagata University School of Medicine, Yamagata, Japan

Yasuharu Imai

Affiliation Department of Internal Medicine, Ikeda Municipal Hospital, Ikeda, Japan

Miki Kokubo

Affiliation Center for Genomic Medicine, Kyoto University Graduate School of Medicine, Kyoto, Japan

Toshihide Shima

Affiliation Center of Gastroenterology and Hepatology, Saiseikai Suita Hospital, Suita, Japan

Hyohun Park

Affiliation Center of Gastroenterology and Hepatology, Saiseikai Suita Hospital, Suita, Japan

Hideo Tanaka

Affiliation Division of Epidemiology and Prevention, Aichi Cancer Center, Nagoya, Japan

Kazuo Tajima

Affiliation Division of Epidemiology and Prevention, Aichi Cancer Center, Nagoya, Japan

Ryo Yamada

Affiliation Center for Genomic Medicine, Kyoto University Graduate School of Medicine, Kyoto, Japan

Fumihiko Matsuda

fumi@genome.med.kyoto-u.ac.jp

Affiliations Center for Genomic Medicine, Kyoto University Graduate School of Medicine, Kyoto, Japan, Institut National de la Sante et de la Recherche Medicale (INSERM) Unite U852, Kyoto University Graduate School of Medicine, Kyoto, Japan

Takeshi Okanoue4 for the Japan Study Group of Nonalcoholic Fatty Liver Disease (JSG-NAFLD)

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FM TO. Performed the experiments: MT M. Kokubo. Analyzed the data: TK RY FM. Contributed reagents/materials/analysis tools: TK YS AU KM MT TT KY T. Saibara EH M. Kokubo SW SK YI M. Kawanaka T. Shima HP HT KT RY. Wrote the paper: TK MT RY FM TO.