About the Authors

Sarah E. Dedonder

Affiliation Department of Diagnostic Medicine/Pathobiology, College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Chuanmin Cheng

Affiliation Department of Diagnostic Medicine/Pathobiology, College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Lloyd H. Willard

Affiliation Department of Diagnostic Medicine/Pathobiology, College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Daniel L. Boyle

Affiliation Division of Biology, College Of Arts And Sciences, Kansas State University, Manhattan, Kansas, United States of America

Roman R. Ganta

rganta@vet.ksu.edu

Affiliation Department of Diagnostic Medicine/Pathobiology, College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Competing Interests

RRG is a PLoS ONE Editorial Board member. This does not alter the authors’ adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RRG. Performed the experiments: SED CC LHW DLB. Analyzed the data: SED RRG. Contributed reagents/materials/analysis tools: SED CC LHW DLB RRG. Wrote the paper: SED RRG.