About the Authors

Brittany N. Sprecher

Affiliation Sinsheimer Laboratories, Department of Molecular, Cellular and Developmental Biology, University of California Santa Cruz, Santa Cruz, California, United States of America

Margo E. Gittings

Affiliation Sinsheimer Laboratories, Department of Molecular, Cellular and Developmental Biology, University of California Santa Cruz, Santa Cruz, California, United States of America

Robert A. Ludwig

rludwig@ucsc.edu

Affiliation Sinsheimer Laboratories, Department of Molecular, Cellular and Developmental Biology, University of California Santa Cruz, Santa Cruz, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RL. Performed the experiments: BS MG RL. Analyzed the data: BS MG RL. Contributed reagents/materials/analysis tools: BS MG RL. Wrote the paper: RL.