About the Authors

Ming Shao

Affiliation Key Laboratory of Experimental Teratology of the Ministry of Education, Key Laboratory of Animal Cells and Developmental Biology of Shandong Province, Life Science College, Shandong University, Jinan, China

Yushuang Lin

Affiliation Key Laboratory of Experimental Teratology of the Ministry of Education, Key Laboratory of Animal Cells and Developmental Biology of Shandong Province, Life Science College, Shandong University, Jinan, China

Zhongzhen Liu

Affiliation Key Laboratory of Experimental Teratology of the Ministry of Education, Key Laboratory of Animal Cells and Developmental Biology of Shandong Province, Life Science College, Shandong University, Jinan, China

Ying Zhang

Affiliation Key Laboratory of Experimental Teratology of the Ministry of Education, Key Laboratory of Animal Cells and Developmental Biology of Shandong Province, Life Science College, Shandong University, Jinan, China

Lifeng Wang

Affiliation Key Laboratory of Experimental Teratology of the Ministry of Education, Key Laboratory of Animal Cells and Developmental Biology of Shandong Province, Life Science College, Shandong University, Jinan, China

Changbin Liu

Affiliation Key Laboratory of Experimental Teratology of the Ministry of Education, Key Laboratory of Animal Cells and Developmental Biology of Shandong Province, Life Science College, Shandong University, Jinan, China

Hongwei Zhang

zhw@sdu.edu.cn

Affiliation Key Laboratory of Experimental Teratology of the Ministry of Education, Key Laboratory of Animal Cells and Developmental Biology of Shandong Province, Life Science College, Shandong University, Jinan, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MS HZ. Performed the experiments: MS YL ZL YZ LW. Analyzed the data: MS YL CL HZ. Wrote the paper: MS HZ.