About the Authors

Satoshi Sugimura

Affiliation National Livestock Breeding Center, Nishigo, Fukushima, Japan

Tomonori Akai

Affiliation Dai Nippon Printing Co., Ltd., Kashiwa, Chiba, Japan

Yutaka Hashiyada

Affiliation National Livestock Breeding Center, Nishigo, Fukushima, Japan

Tamás Somfai

Affiliations National Livestock Breeding Center, Nishigo, Fukushima, Japan, National Institute of Livestock and Grassland Science, Tsukuba, Ibaraki, Japan

Yasushi Inaba

Affiliations National Livestock Breeding Center, Nishigo, Fukushima, Japan, National Institute of Livestock and Grassland Science, Tsukuba, Ibaraki, Japan

Muneyuki Hirayama

Affiliation National Livestock Breeding Center, Nishigo, Fukushima, Japan

Tadayuki Yamanouchi

Affiliation National Livestock Breeding Center, Nishigo, Fukushima, Japan

Hideo Matsuda

Affiliation National Livestock Breeding Center, Nishigo, Fukushima, Japan

Shuji Kobayashi

Affiliation National Livestock Breeding Center, Nishigo, Fukushima, Japan

Yoshio Aikawa

Affiliation National Livestock Breeding Center, Nishigo, Fukushima, Japan

Masaki Ohtake

Affiliation National Livestock Breeding Center, Nishigo, Fukushima, Japan

Eiji Kobayashi

Affiliation National Livestock Breeding Center, Nishigo, Fukushima, Japan

Kazuyuki Konishi

Affiliation National Livestock Breeding Center, Nishigo, Fukushima, Japan

Kei Imai

k0imai@nlbc.go.jp

Affiliation National Livestock Breeding Center, Nishigo, Fukushima, Japan

Competing Interests

Cell culture dish (No. 4724854) was co-applied by Dai Nippon Printing Co., Ltd., and National Livestock Breeding Center. ID culture is produced and marketed by Dai Nippon Printing Co., Ltd. TA is employed by Dai Nippon Printing Co., Ltd. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SS KI. Performed the experiments: SS TA YH TS YI MH TY HM SK YA MO EK KK KI. Analyzed the data: SS. Contributed reagents/materials/analysis tools: SS TA KI. Wrote the paper: SS.