About the Authors

Tomoaki Nishiyama

Contributed equally to this work with: Tomoaki Nishiyama, Kaori Miyawaki

tomoakin@staff.kanazawa-u.ac.jp (TN); tekurata@bs.naist.jp (TK); mhasebe@nibb.ac.jp (MH)

Affiliations Exploratory Research for Advanced Technology, Japan Science and Technology Agency, Okazaki, Japan, Advanced Science Research Center, Kanazawa University, Kanazawa, Japan

Kaori Miyawaki

Contributed equally to this work with: Tomoaki Nishiyama, Kaori Miyawaki

Current address: Department of Botany and Plant Sciences, Center for Plant Cell Biology, University of California Riverside, Riverside, California, United States of America

Affiliation Exploratory Research for Advanced Technology, Japan Science and Technology Agency, Okazaki, Japan

Masumi Ohshima

Affiliation Exploratory Research for Advanced Technology, Japan Science and Technology Agency, Okazaki, Japan

Kari Thompson

Affiliation Exploratory Research for Advanced Technology, Japan Science and Technology Agency, Okazaki, Japan

Akitomo Nagashima

Affiliation Exploratory Research for Advanced Technology, Japan Science and Technology Agency, Okazaki, Japan

Mitsuyasu Hasebe

tomoakin@staff.kanazawa-u.ac.jp (TN); tekurata@bs.naist.jp (TK); mhasebe@nibb.ac.jp (MH)

Affiliations Exploratory Research for Advanced Technology, Japan Science and Technology Agency, Okazaki, Japan, Division of Evolutionary Biology, National Institute for Basic Biology, Okazaki, Japan, School of Life Science, The Graduate University for Advanced Studies, Okazaki, Japan

Tetsuya Kurata

tomoakin@staff.kanazawa-u.ac.jp (TN); tekurata@bs.naist.jp (TK); mhasebe@nibb.ac.jp (MH)

Current address: Plant Global Education Project, Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma, Japan

Affiliation Exploratory Research for Advanced Technology, Japan Science and Technology Agency, Okazaki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TK KM TN MH. Performed the experiments: KM TN MO KT AN MH TK. Analyzed the data: KM TN MH TK. Wrote the paper: TN KM TK MH.