About the Authors

Fumihiro Niwa

Affiliations Laboratory for Developmental Neurobiology, Brain Science Institute (BSI), RIKEN, Saitama, Japan, Laboratory of Functional Genomics, Department of Medical Genome Science, Graduate School of Frontier Science, The Institute of Medical Science, The University of Tokyo, Minato-ku, Tokyo, Japan

Hiroko Bannai

hbannai@brain.riken.jp

Affiliation Laboratory for Developmental Neurobiology, Brain Science Institute (BSI), RIKEN, Saitama, Japan

Misa Arizono

Affiliations Laboratory for Developmental Neurobiology, Brain Science Institute (BSI), RIKEN, Saitama, Japan, Division of Neuronal Network, The Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Kazumi Fukatsu

Affiliation Laboratory for Developmental Neurobiology, Brain Science Institute (BSI), RIKEN, Saitama, Japan

Antoine Triller

Affiliation Institut de Biologie de l'École Normale Supérieure (IBENS), Institut National de la Santé et de la Recherche Médicale U1024, Centre National de la Recherche Scientifique UMR8197, Paris, France

Katsuhiko Mikoshiba

Affiliation Laboratory for Developmental Neurobiology, Brain Science Institute (BSI), RIKEN, Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HB AT KM. Performed the experiments: FN HB MA. Analyzed the data: FN HB. Contributed reagents/materials/analysis tools: FN HB KF. Wrote the paper: FN HB MA AT KM.