About the Authors

Ho-Hsing Tseng

Contributed equally to this work with: Ho-Hsing Tseng, Qiu-Yu Chuah

Affiliation Department of Life Science, Tzu Chi University, Hualien, Taiwan R.O.C.

Qiu-Yu Chuah

Contributed equally to this work with: Ho-Hsing Tseng, Qiu-Yu Chuah

Affiliation Department of Life Science, Tzu Chi University, Hualien, Taiwan R.O.C.

Pei-Ming Yang

Affiliation Department of Pharmacology, College of Medicine, National Taiwan University, Taipei, Taiwan R.O.C.

Chiung-Tong Chen

Affiliation Institute of Biotechnology and Pharmaceutical Research, National Health Research Institutes, Zhunan, Taiwan R.O.C.

Jung-Chi Chao

Affiliation Department of Life Science, Tzu Chi University, Hualien, Taiwan R.O.C.

Ming-Der Lin

Affiliation Department of Molecular Biology and Human Genetic, Tzu Chi University, Hualien, Taiwan R.O.C.

Shu-Jun Chiu

chiusj@mail.tcu.edu.tw

Affiliation Department of Life Science, Tzu Chi University, Hualien, Taiwan R.O.C.

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: S-JC. Performed the experiments: H-HT Q-YC J-CC. Analyzed the data: S-JC P-MY. Contributed reagents/materials/analysis tools: C-TC M-DL. Wrote the paper: S-JC P-MY H-HT.