About the Authors

Milla Summanen

Affiliations Cell Biology, Department of Biology, University of Utrecht, Utrecht, The Netherlands, Division of Pharmacology and Toxicology, Faculty of Pharmacy, University of Helsinki, Helsinki, Finland

Niko Granqvist

Affiliation Division of Biopharmaceutics and Pharmacokinetics, Faculty of Pharmacy, University of Helsinki, Helsinki, Finland

Raimo K. Tuominen

Affiliation Division of Pharmacology and Toxicology, Faculty of Pharmacy, University of Helsinki, Helsinki, Finland

Marjo Yliperttula

Affiliation Division of Biopharmaceutics and Pharmacokinetics, Faculty of Pharmacy, University of Helsinki, Helsinki, Finland

C. Theo Verrips

Affiliation Cell Biology, Department of Biology, University of Utrecht, Utrecht, The Netherlands

Johannes Boonstra

j.boonstra@uu.nl

Affiliation Cell Biology, Department of Biology, University of Utrecht, Utrecht, The Netherlands

Christophe Blanchetot

Contributed equally to this work with: Christophe Blanchetot, Elina Ekokoski

Affiliation Cell Biology, Department of Biology, University of Utrecht, Utrecht, The Netherlands

Elina Ekokoski

Contributed equally to this work with: Christophe Blanchetot, Elina Ekokoski

Affiliation Division of Pharmacology and Toxicology, Faculty of Pharmacy, University of Helsinki, Helsinki, Finland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MS NG CB EE. Performed the experiments: MS NG. Analyzed the data: MS NG. Contributed reagents/materials/analysis tools: MY CTV. Wrote the paper: MS NG RKT JB CB EE.