About the Authors

Sophie Richier

s.richier@noc.soton.ac.uk

Affiliation Ocean and Earth Science, National Oceanography Centre, Southampton, University of Southampton, European Way, Southampton, United Kingdom

Anna I. Macey

Affiliation Ocean and Earth Science, National Oceanography Centre, Southampton, University of Southampton, European Way, Southampton, United Kingdom

Nicola J. Pratt

Affiliation Ocean and Earth Science, National Oceanography Centre, Southampton, University of Southampton, European Way, Southampton, United Kingdom

David J. Honey

Affiliation Ocean and Earth Science, National Oceanography Centre, Southampton, University of Southampton, European Way, Southampton, United Kingdom

C. Mark Moore

Affiliation Ocean and Earth Science, National Oceanography Centre, Southampton, University of Southampton, European Way, Southampton, United Kingdom

Thomas S. Bibby

Affiliation Ocean and Earth Science, National Oceanography Centre, Southampton, University of Southampton, European Way, Southampton, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SR TSB CMM. Performed the experiments: SR NJP DJH. Analyzed the data: SR NJP CMM TSB. Contributed reagents/materials/analysis tools: SR AIM DJH CMM TSB. Wrote the paper: SR CMM TSB.